The taxonomy of scorpions deals with the classification of this predatory arthropod into 13 extant families and about 1,400 described species and subspecies. In addition, 111 described taxa of extinct scorpions are known.[1]
The classification is based on that of Soleglad and Fet (2003),[2] which replaced the older, unpublished classification of Stockwell.[3] Additional taxonomic changes are from papers by Soleglad et al. (2005).[4][5] The classification proposed by Fet and Soleglad in 2003 and subsequent papers has not been universally accepted; some authorities have challenged their methodology as invalid.[5]
Taxonomy
Order Scorpiones
- Infraorder Orthosterni  Pocock, 1911 
- Parvorder Pseudochactida  Soleglad et Fet, 2003 
- Superfamily Pseudochactoidea  Gromov, 1998 
- Family Pseudochactidae Gromov, 1998
 
 
- Superfamily Pseudochactoidea  Gromov, 1998 
- Parvorder Buthida  Soleglad et Fet, 2003 
- Superfamily Buthoidea  C. L. Koch, 1837 
- Family Buthidae C. L. Koch, 1837 (thick-tailed scorpions)
- Family Microcharmidae Lourenço, 1996
 
 
- Superfamily Buthoidea  C. L. Koch, 1837 
- Parvorder Chaerilida  Soleglad et Fet, 2003 
- Superfamily Chaeriloidea  Pocock, 1893 
- Family Chaerilidae  Pocock, 1893 
- Subfamily Chearilinae Pocock, 1893
- † Subfamily Electrochearilinae Fet, Soleglad et Anderson 2004 (extinct)
 
 
- Family Chaerilidae  Pocock, 1893 
 
- Superfamily Chaeriloidea  Pocock, 1893 
- Parvorder Iurida  Soleglad et Fet, 2003 
- Superfamily Chactoidea  Pocock, 1893 
- Family Chactidae Pocock, 1893
- Family Euscorpiidae  Laurie, 1896 
- Subfamily Euscorpiinae Laurie, 1896
- Subfamily Megacorminae  Kraepelin, 1905 
- Tribe Chactopsini Soleglad et Sissom, 2001
- Tribe Megacormini Kraepelin, 1905
 
- Subfamily Scorpiopinae  Kraepelin, 1905 
- Tribe Scorpiopini Kraepelin, 1905
- Tribe Troglocormini Soleglad et Sissom, 2001
 
 
- Family Superstitioniidae Stahnke, 1940
- Family Vaejovidae Thorell, 1876
 
- Superfamily Iuroidea  Thorell, 1876 
- Family Caraboctonidae  Kraepelin, 1905 (hairy scorpions)
- Subfamily Caraboctoninae Kraepelin, 1905
- Subfamily Hadrurinae Stahnke, 1974
 
- Family Iuridae Thorell, 1876
 
- Family Caraboctonidae  Kraepelin, 1905 (hairy scorpions)
- Superfamily Scorpionoidea  Latreille, 1802 
- Family Bothriuridae  Simon, 1880 
- Subfamily Bothriurinae Simon, 1880
- Subfamily Lisposominae Lawrence, 1928
 
- Family Hemiscorpiidae Pocock, 1893 (= Ischnuridae, =Liochelidae) (rock scorpions, creeping scorpions, or tree scorpions)
- † Family Protoischnuridae Carvalho et Lourenço 2001 (extinct)
- Family Scorpionidae Latreille, 1802 (burrowing scorpions or pale-legged scorpions)
 
- Family Bothriuridae  Simon, 1880 
- Superfamily incertae sedis
- † Family Palaeoeuscorpiidae Lourenço 2003 (extinct)
 
 
- Superfamily Chactoidea  Pocock, 1893 
- Parvorder incertae sedis
- † Family Archaeobuthidae Lourenço 2003 (extinct)
- † Family Palaeopisthacanthidae Kjellesvig-Wæring 1986 (extinct)
- † Family Protobuthidae Lourenço et Gall 2003 (extinct)
- † Family incertae sedis (contains genera Corniops Jeram 1994; Palaeoburmesebuthus Lourenço 2002; Sinoscorpius Hong 1983).
 
 
- Parvorder Pseudochactida  Soleglad et Fet, 2003 
References
- ↑ Dunlop, Jason A.; Penney, David; Tetlie, O. Erik; Anderson, Lyall I. (2008). "How many species of fossil arachnids are there". Journal of Arachnology. BioOne. 36 (2): 262–272. doi:10.1636/CH07-89.1. Retrieved 2010-04-07.
- ↑ Soleglad, Michael E.; Fet, Victor (2003). "High-level systematics and phylogeny of the extant scorpions (Scorpiones: Orthosterni)" (multiple parts). Euscorpius. Marshall University. 11: 1–175. Retrieved 2008-06-13.
- ↑ Stockwell, Scott A., 1989. Revision of the Phylogeny and Higher Classification of Scorpions (Chelicerata). Ph.D. Dissertation, University of California, Berkeley
- ↑ Soleglad, Michael E.; Fet, Victor; Kovařík, F. (2005). "The systematic position of the scorpion genera Heteroscorpion Birula, 1903 and Urodacus Peters, 1861 (Scorpiones: Scorpionoidea)" (PDF). Euscorpius. Marshall University. 20: 1–38. Retrieved 2008-06-13.
- 1 2 Fet, V.; Soleglad, Michael E. (2005). "Contributions to scorpion systematics. I. On recent changes in high-level taxonomy" (PDF). Euscorpius. Marshall University (31): 1–13. ISSN 1536-9307. Retrieved 2010-04-07.
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